Description

D e s c r i p t i o n a n d r e m a r k s. Goczania rugosa is represented in the Catefica mesofossil flora by two anthers and an isolated pollen sac with pollen grains in situ (Textfig. 20 e – h). Identical pollen also occurs in pollen clumps and coprolites. The anther is short and broad, about 0.6 mm long and 0.55 mm wide, dithecate and tetrasporangiate (Text-fig. 20 e). As in the type material from Torres Vedras, the inner wall of the anthers of the Catefica specimen and the in situ pollen grains show numerous small, spherical orbicules with a finely spiny surface ornamentation (Text-fig. 20 f). The pollen grains are oblate, circular to elliptical in equatorial outline, about 17 µm in diameter and monocolpate (Text-fig. 20 f – h). The colpus is short with an irregular margin (Text-fig. 20 h). The exine is tectate with the tectum covered with densely spaced microechinae that occur singly without merging with their neighbors (Text-fig. 20 f – h). A f f i n i t y a n d o t h e r o c c u r r e n c e s. Pollen of the Goczania type has been found on the stigma and surface of Appomattoxia fruits in the Torres Vedras mesofossil flora and also on fruits of Appofructus E. M. FRIIS, P. R. CRANE et K. R. PEDERSEN from Torres Vedras (Friis et al. 2019 a). Goczania- type pollen has also been found on the stigma and surface of Appomattoxia fruits from the Puddledock mesofossil flora of eastern North America. Appomattoxia and Appofructus are both thought to be related to Piperales (Friis et al. 1995, 2019 a).

Common Names

Taxonomic Hierarchy

Kingdom: Plantae

Phylum: Tracheophyta

Class: Magnoliopsida

Order: Chloranthales

Family: Chloranthaceae

Genus: Goczania

Species: Goczania rugosa E.M.Friis, P.R.Crane & K.R.Pedersen, 2019